New Osmylopsychopid Taxa from the Middle Jurassic of Northeastern China (Neuroptera: Osmylopsychopidae) †

SIMPLE SUMMARY: Osmylopsychopidae, an extinct family of Neuroptera, is closely related to the extant Psychopsidae. The sporadic records of this family and limited preservation of fossil specimens have hindered the understanding of the evolution of Osmylopsychopidae. In this paper, one new genus with four new species of Osmylopsychopidae are described from the Middle Jurassic Jiulongshan Formation of northeastern China. These newly discovered osmylopsychopids with a forewing length of ca. 10 mm (implying small body size), clearly distinguished from the traditional sense of Osmylopsychopidae, suggest that Osmylopsychopidae possibly underwent a significant diversification during the Mesozoic. ABSTRACT: One new genus with four new species sharing the similar character of a forewing length of ca. 10 mm, namely, Minipsychops spasulus gen. et sp. nov., Minipsychops polychotomus sp. nov., Minipsychops densus sp. nov., and Minipsychops unicus sp. nov., are described from the Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia in China. These new insects can be attributed to Osmylopsychopidae for their distinctive configuration of costal space and the venations of RP1 and Cu. Compared with the typically medium to large body sizes of known osmylopsychopids, these new Middle Jurassic taxa of a particularly miniaturized lineage not only enrich the species diversity of Osmylopsychopidae, but also enhance our understanding of the historical evolution of these poorly known lacewings.