A revised circumscription for the Blakeeae (Melastomataceae) with associated nomenclatural adjustments

Abstract. Systematic investigations and phylogenetic analyses of the Blakeeae (Melastomataceae) have indicated that Topobea should be synonymized under Blakea, and Huilaea under Chalybea. Presented here is a detailed description of the Blakeeae, a key to its two accepted genera, and a listing of 62 new combinations, including 3 new names, necessitated by the transfer of Topobea as follows: Blakea acuminata (Wurdack) Penneys & Judd, comb. nov., Blakea adscendens (E.Cotton & Matezki) Penneys & Judd, comb. nov., Blakea albertiae (Wurdack) Penneys & Almeda, comb. nov., Blakea amplifolia (Almeda) Penneys & Almeda, comb. nov., Blakea arboricola (Almeda) Penneys & Almeda, comb. nov., Blakea asplundii (Wurdack) Penneys & Judd, comb. nov., Blakea barbata (Gleason) Penneys & Judd, comb. nov., Blakea brenesii (Standl.) Penneys & Almeda, comb. nov., Blakea brevibractea (Gleason) Penneys & Judd, comb. nov., Blakea bullata (E.Cotton & Matezki) Penneys & Judd, comb. nov., Blakea calcarata (L.Uribe) Penneys & Judd, comb. nov., Blakea calophylla (Almeda) Penneys & Almeda, comb. nov., Blakea calycularis (Naudin) Penneys & Almeda, comb. nov., Blakea castanedae (Wurdack) Penneys & Judd, comb. nov., Blakea clavata (Triana) Penneys & Judd, nom. nov., Blakea cordata (Gleason) Penneys & Almeda, comb. nov., Blakea cuprina Penneys & Judd, nom. nov., Blakea cutucuensis (Wurdack) Penneys & Judd, comb. nov., Blakea dimorphophylla (Almeda) Penneys & Almeda, comb. nov., Blakea discolor (Hochr.) Penneys & Judd, comb. nov., Blakea dodsonorum (Wurdack) Penneys & Almeda, comb. nov., Blakea eplingii (Wurdack) Penneys & Judd, comb. nov., Blakea ferruginea (Gleason) Penneys & Judd, comb. nov., Blakea fragrantissima (Almeda) Penneys & Almeda, comb. nov., Blakea gerardoana (Almeda) Penneys & Almeda, comb. nov., Blakea glaberrima (Triana) Penneys & Judd, comb. nov., Blakea henripittieri (Cogn.) Penneys & Almeda, comb. et nom. nov., Blakea hexandra (Almeda) Penneys & Almeda, comb. nov., Blakea horologica Penneys & Judd, nom. nov., Blakea induta (Markgr.) Penneys & Judd, comb. nov., Blakea inflata (Triana) Penneys & Judd, comb. nov., Blakea insignis (Triana) Penneys & Judd, comb. nov., Blakea intricata (Almeda) Penneys & Almeda, comb. nov., Blakea killipii (Wurdack) Penneys & Judd, comb. nov., Blakea lentii (Almeda) Penneys & Almeda, comb. nov., Blakea longiloba (Wurdack) Penneys & Judd, comb. nov., Blakea longisepala (Gleason) Penneys & Judd, comb. nov., Blakea macbrydei (Wurdack) Penneys & Judd, comb. nov., Blakea maguirei (Wurdack) Penneys & Judd, comb. nov., Blakea maurofernandeziana (Cogn.) Penneys & Almeda, comb. nov., Blakea mcphersonii (Almeda) Penneys & Almeda, comb. nov., Blakea modica (Wurdack) Penneys & Judd, comb. nov., Blakea mortoniana (Wurdack) Penneys & Judd, comb. nov., Blakea muricata (Lozano) Penneys & Judd, comb. nov., Blakea pascoensis (Wurdack) Penneys & Judd, comb. nov., Blakea pluvialis (Standl.) Penneys & Almeda, comb. nov., Blakea sessilifolia (Triana) Penneys & Judd, comb. nov., Blakea setosa (Triana) Penneys & Judd, comb. nov., Blakea standleyi (L.O.Williams) Penneys & Almeda, comb. nov., Blakea stephanochaeta (Naudin) Penneys & Judd, comb. nov., Blakea steyermarkii (Wurdack) Penneys & Judd, comb. nov., Blakea suaveolens (Almeda) Penneys & Almeda, comb. nov., Blakea subbarbata (Wurdack) Penneys & Judd, comb. nov., Blakea subscabrula (Triana) Penneys & Judd, comb. nov., Blakea subsessiliflora (Wurdack) Penneys & Judd, comb. nov., Blakea superba (Naudin) Penneys & Judd, comb. nov., Blakea tetramera (Almeda) Penneys & Almeda, comb. nov., Blakea tetroici (Wurdack) Penneys & Judd, comb. nov., Blakea toachiensis (Wurdack) Penneys & Judd, comb. nov., Blakea trianae (Cogn.) Penneys & Judd, comb. nov., Blakea verrucosa (Wurdack) Penneys & Judd, comb. nov., Blakea watsonii (Cogn.) Penneys & Almeda, comb. nov.