The genus Scaptodrosophila Duda part I: the brunnea species group from the Oriental Region, with morphological and molecular evidence (Diptera, Drosophilidae)

Abstract. Seven new species of the Scaptodrosophila brunnea species group are described from east Asia: S. maculata sp. n., S. melanogaster sp. n., S. nigricostata sp. n., S. nigripecta sp. n., S. obscurata sp. n., S. protenipenis sp. n. and S. rhina sp. n. Three known species, S. parabrunnea (Tsacas & Chassagnard), S. pressobrunnea (Tsacas & Chassagnard) and S. scutellimargo (Duda) are redescribed. A key to all the examined species in the brunnea group is provided. Species delimitations have been improved by integrating the DNA sequences with morphological information. The intra- and interspecific pairwise p-distances (proportional distance) are summarized. Some nucleotide sites with fixed status in the alignment of the COI sequences (664 nucleotide sites in length) are used as “pure” molecular diagnostic characters to delineate species in the brunnea group.