Recent Advances in Phytohormone Regulation of Apple-Fruit Ripening

Apple (Malus domestica) is, globally, one of the largest fruits in terms of cultivated area and yield. Apple fruit is generally marketed after storage, which is of great significance for regulating the market supply in the off-season of fruit production. Apple-fruit ripening, which culminates in desirable changes in structural and textural properties, is governed by a complex regulatory network. Much is known about ethylene as one of the most important factors promoting apple-fruit ripening. However, the dynamic interplay between phytohormones also plays an important part in apple-fruit ripening. Here, we review and evaluate the complex regulatory network concerning the action of phytohormones during apple-fruit ripening. Interesting future research areas are discussed.