Two new pendulous epiphytic Columnea L. (Gesneriaceae) species from the Chocó forests of the Northern Andes

Exploratory field expeditions to the Chocó forests in the northwestern slopes of the Ecuadorian and Colombian Andes resulted in the discovery of two new species of Columnea (Gesneriaceae). Columneafluidifolia J.L.Clark & F.Tobar, sp. nov., is described as a narrow endemic from Bosque Protector Mashpi and surrounding areas in the province of Pichincha in northern Ecuador. Columneapendens F.Tobar, J.L.Clark & J.F.Sm., sp. nov., is described from recently discovered populations in the provinces of Carchi and Santo Domingo de los Tsáchilas (Ecuador) and the departments of Cauca and Nariño in southwestern Colombia. The two new species are pendent epiphytes with elongate shoots and shallowly bilabiate to nearly tubular corollas. Descriptions, complete specimen citations, and a distribution map are provided. Based on IUCN guidelines, a preliminary conservation status of Critically Endangered (CR) is provided for C.fluidifolia and Endangered (EN) is provided for C.pendens.